Light strikes photosystem-2 by exciting the electrons. The electron comes from a water molecule that has been split into hydrogen and oxygen. The excited electrons then travel down a short electron transport chain, and while doing so, they pump H+ into the inner thylakoid space across the thylakoid membrane from the stroma. The electrons are then passed, at the end of the transport chain, to photosystem 1, where they are re-excited by light.