Actinostele is the modification of
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Protostele
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Siphonostele
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Solenostele
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Dictyostele
The earliest vascular plants had stems with a central core of vascular tissue. This consisted of a cylindrical strand of xylem, surrounded by a region of phloem. Around the vascular tissue, there might have been an endodermis that regulated the flow of water into and out of the vascular system. Such, an arrangement is termed a protostele, which is the simplest type of stele. Actinostele is a variation of the protostele in which the core is lobed or fluted. This stele is found in many species of club moss. Actinosteles are typically exarch and consist of several to many patches of protoxylem at the tips of the lobes of the metaxylem. Exarch protosteles are a defining characteristic of the lycophyte lineage.