Which of the following options is the best substitute for the underlined words in statement III?

Directions: In this passage, errors or parts of a sentence, which have been left out, have been underlined. To correct these errors and correctly complete the passage, choose the correct option.
Statement I: In India, there stood a colony of crows on a huge tree, known
Statement II: as banyan tree. On many tree nearby, lived a colony of owls.
Statement III: The owls is powerful enemies, frequently attacking and harassing
Statement IV: the crows. The owls could see at night, _________.