In a rigid flanged coupling, bolts are primarily loaded in shear due to the torque transmitted between shafts. The rigid connection also creates bending moments in the bolts because the flanges cannot rotate relative to each other, causing the bolts to resist both shear and bending. Axial force alone (B) is incorrect because the primary load is not along the bolt axis. Pure torsion (C) doesn't apply to bolts in this configuration. While torsion exists in the shafts, the bolts themselves experience shear and bending, not shaft torsion.